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It is difficult to know exactly when the shift from gelatine to agar occurred. As often happens for scientific breakthroughs, agar was likely adopted incrementally alongside the use of other growth media. In 1913, for example, the first diagnosis of Serratia marcescens as a human pathogen was made by growing it on agar as well as on potatoes.
Ac we not free ne sindon, for-thy-the we never ne mighton from Wulfsfleet yewitan, neven we thone Hlaford finden and hine ofslain. Se Hlaford has thisne stede mid searocraftum yebounden, that none ne may hine forletan. We sindon here like fuglas in a nete, like fishes in a weir.